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(1936) [MARC] Author: Rutger Sernander
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liypothesis is founded on a sa-ious undeiestimation of ti» length of time
needed by a spiuce to giosv fi-orn oxigo to bi-east lieiglit An aiov of a
guai-ta- of a cesstuiy must, bo \?e vex, not be undaiated is a discussion of this
bind. Tbe valnes obtai’/»d by tlie piesent authox show tliat, in spite of
xavaging stoims dming tlie 13th and 2Ötli centm-ies, spiuce; fiom yeais befove
173ö stand close enough to malte it isnpossible to suppose tliat tlie foiest has
developed fiom a pastnie, atandoned at tlie dis continuatio n of tiie pstteiy
factoi-y in 17fi Tx.

Thus, tbe eoology of tlie dviaif tiee is one of tbe impoxtant factois in
an illucidation of tlie liistoiy of tbe stoxm-gap foiest It is obvions tliat tlie
bis t development of tlie spiuce is las snccessful in tlie noinial bottom
laya-than on nedaphidesn. Socli "edaplsides» aie exeinplified by oveitnmed
l-oot-systesris, laige boulda-s and fallen tinnlcs (Swedish »låga», pi. ulSgpii).
Bstamples aie to be found on p. 1(13 167.

Tbe decaying fallen tinnlis play an impsitant idle in tlie liista-y of tlie
s pmce foiest, and tlie autliox bas tlieiefoie dxaivn up a scale nneaotisation«
iritli 6 degiees. Gaminating seeds of tlie spiuce aie found along tinnlrs in
stages 5 it 6 of neciotisation. As in tlie elassical piotuies fi-om Kubany by
Gonso-ÄX, it is also easy at Giansltäx and in Ti by to find geisainating young
spyuoes is io\rs because of tlie leg-enexatian along tbe decaying fallen tivnlcs.
An instiuctive pictuie of sucli a iqw of tiees about 130 yeais old is to be
seen on fig. 03. Of tbe fallen tiunlt no lemains can te obseived, and
afta-tlie laps of absut SKJ yeais afta- tbe fall of a taunlt it seessis to be
impossible to detect iti piesence in tlie covex of Äpicvwim; ji; , tbe ooIonisation
of ivliicli indicates neaotisation degi-ee 6.

In lefeience to tlie oindi tions in tlie eutiopbic paits of tbe Ku ta ny
pi-imeval sptuce foi-ets, it lias teen claimed tliat ttie noinial type of taancliing
and an ef 11 den t cleaiing of tvrigs (comp. tig. 63) sliould chai-acteiize all spi-uces is
a natuial foiest Sudi a gaseialisation is, liowevei-, nnwai-ianted. It doa not
apply to tlie oligoti-ophic Iify(o.w>iiifUj-spiuce-foi-est of Tiby; teie tlie spiuces
may te ii-iegulai-ly bi-anclied (nspängienighetu, comp. fig. (JO), bnt in mai-e
eutiopbic, lei-baceous, paits tliey become noxmaliy bianclied, comp. figs. JiO öt.

At tlie fii-st visit to a natm-al oi- pi-isneval spiuce foiest, one may be
sux-pi-ised to find tliat tlie Unnks aie not so laige as one v/oold net-haps have
eepeeted. Hiis physionomical featui-e is ds» to two factois: 1) tl» small annual
diameta- gi-owtli wlia-e tbe vitality of tlie spiuce has been iedvoed by density
of tlie staisi oi- by i-oot fnngi so tliat tlie tiunlis appeai- yovosga- tlian
tliey leally aie, and 2) tbe pai-ticulax susceptibility of tlie spi-nces to
stoi-m-feUiisg. Tbis latta- factox, vdiicb bas eai-liei- teen neglected oi- misnndeistood,
desei-vss special empliasis. Tlie fact ttiat tlie foiest looks os mpaiati sely young
is ivell-illustxated by a gieat numba- of boisngs, made by Hlissmosaw in
oi-dei- to pio>e a conti-ai-y contesssion in a numbei- of spiuces gi-ovong up

into tlie ci-owns of laige biiebes and aspens. Tlese bii-clies piove to te

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