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SVENSK BOTANISK TIDSKRIFT. 1921: Bb. 15, H. 1.

ECOLOGICAL STUDIES IN THE ASSIMILATION
OF CERTAIN FOREST-PLANTS AND
SHORE-PLANTS.

BY
HENRIK LUNDEGARDH.

CONTRIBUTIONS FROM HALLANDS VADERO ECOLOGICAL STATION. N:o4

1. Introduction.

In the study of the carbon dioxide assimilation the attention has
been directed partly upon the investigation of the internal conditions
and the chemical nature of the process, and partly upon the de-
duction of the external factors affecting the course of the assimilation.
Among the most important of the chemical works dealing with the
carbon dioxide assimilation must undoubledly be mentioned the
researches of WILLSTATTER and STOLL (1913, 1918). Our knowledge
of the external conditions has been carried a great step forwards
more particularly by the labours of BLACKMAN and his collabo-
rators.

According to our present knowledge the assimilation is subject
to the following conditions: —1, the amount of chlorophyll and 2,
a protoplasm factor (the »assimilation enzym» of WILLSTATTER and
Srorr [1918], which together form the internal factors, and 3, the
amount of carbon dioxide, 4, the amount of water, 5, the supply
of light, and 6, the temperature, which together make up the external
factors.

In regard to the reciprocal influence of the various external factors
the theory put forward by BLACKMAN seems to be generally adop-
ted. According to BLackMAN (1905, BLACKMAN and MATTHAEI 1905,
Brackman and SMITH 1911), the factor present in the minimum

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