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372
HENNING HORN AF RANTZIEN
From what precedes it will be seen that 110 reasons can be found
for the arrangement of Phl. arenarium and related forms by Fiori
and Paoletti (1896—98) and Fiori (1923) where Phl. arenarium ssp.
eu-arenarium, Piil. crypsoides ssp. sardoum and Phl. exaratum are
iinited into one species (ef. Vierhapper I.e., p. 306). It will also be seen
that Phl. arenarium phvtogeographically holds a verv interesting special
position among the related forms within Chilochloa. having its centre
of distribution in eu-oceanic Western Europé and not as the other
mentioned species in the east of the Mediterranean Area, with the
clearly accentuated summer drought of these districts.
In most cases Phl. arenarium has been regarded in the literature
as a European Atlantic — Mediterranean species. However, there are
some statements from other continents too.
In the herb, of Swartz (now preserved in Riksmuseum,
Stockholm) there is a collection labelled »Ex America Boreali seq. Swartz».
All other records from this continent are lacking and we must consider
this statement to be dependent upon a confusion of labels.
Grisebach (1868, p. 83) gives a record of Pld. arenarium from
NW. Himalaya, leg. Thomson. He was of the opinion that this
Hima-layan Phl. arenarium was to be treated as a separate variety, var.
Thomsonii Griseb. Hooker (1897. p. 237) stated a wide distribution
area for this »Phleum arenarium» from the Punjab and W. Himalaya
up to 9.000 feet. hut wrote that he coidd »find no charader whereby
to distinguish this (v. Thomsonii) from the common form».
In 1921 Mez described the Himalayan form as a new species, Phl.
himalaicum Mez (Mez 1921, p. 293). The description is somewhat
incomplete. This species, however, seems to be rather well separated
from Phl. arenarium. The material that has been seen (NW. Himalaya,
alt. 6—8,000 feet, leg. Thomson, preserved in Riksmuseum,
Stockholm. in Mus. Copenhagen and in herb. Delessert and Barbey-Boissier,
Geneva) differs from Phl. arenarium above all in the following characters.
Piil. himalaicum Mez
Laminae of npper leaves ± long and
narrow, often longer than their
vaginae, these being generali}’
shorter than the corresponding
internodes.
Spike tax, especially at the broad,
basal part; often ± paniculate, +
distinctly lobed, also in erect state.
Phl. arenarium L.
Laminae of upper leaves short and
broad, most eommonly shorter
than their vaginae, these generally
being longer than the
corresponding internodes.
Spike dense, compressed, narrowed
towards the basal part, cylindrical,
in erect state seldom lobed.
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