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the female gametophyte in nolana
381
Acantluiceae-forms have both kinds of haustoria and the very
endosperm is nuclear. The same course of development is met with in both
families. 1 conclude that in Acantluiceae the endosperm in Crossandra
and Acanthis is the primitive type, that the rest of species examined by
Mauritzon possess more progressive endosperm forms and that the
endosperm in this family, too, may be derived from a Verbancum-[ype.
Thus it appears that my interpretation of this subtile thing, the
synergid-shaped endosperm cells in Nolana and Hyoscyamus, elucidates
the course of endosperm development in Tubi/’lorae and, as has been
repeatedly emphazised in this paper, leads to the conclusion, that the
origin of the endosperm in all families of this group — the endosperm
development of which is known — most probably can be traced to the
Verbascum-type, that haustoria are primitive characters and the nuclear
endosperm type is the most progressive type in this group.
Botanical institute, Göteborg, September 1947.
Bibliography.
1. Dahlgren, o. Die Embrvologie <ler Loganiceen-Gattung Spigelia. — sv. Bof.
Tidskr. 1(5, 11)22.
2. - Notes on ah initio cellular endosperm. — Bot. Notiser, p. 1—21. 11)23.
3. Glisic, Lj. M. Zur Entwicklungsgeschichte der Solanaceen. Die Endospermbil-
dung von Datura Melel L. -— Bull, de t’inst. el du Jard. botan. de l’univ.
de Beograd. Tome I. p. 75—85. 1928—30.
4. — Ein Versuch der Vervvertung der Endospermmerkmale fiir typologisehe
und phylogenetische Zwecke innerhalb der Scrophulariaceae. Ibid. Tome
IV, p. 42—73. 193(5—1937.
5. Håkansson, A. Zur Zvtologie von Celsia und Verbascum. — Lunds Univ. Arsskr.
N.F. Avd. 2, 21 nr 10. 1926.
6. Kausik, S. B. and Subramanyam, K. A contribution to the life-history of Sphe-
noclea zeylanica Gaertn. — Proc. of Indian Acad, of Sci. Vol. Will. 1946.
7. i.lndl.ey. J. Bol. Reg. 10. Text accompanying t. 865. 1824.
8. Mairitzon, J. Die Endosperm und Embryo-Entwicklung einiger Acanthaceen.
- Lunds Univ. Arsskr. N.F. Avd. 2, Bd 30 nr 5. 1934.
9. Persidsky. J). On Ihe development of endosperm in Solanaceae. - Journ. de
l’inst. bot. de I’acad. des sci. d l’kraine nr 4 (12). p. 35—45. 1935.
10. ROSÉN, \V. Zur Embrvologie der Campanulaceen. — Acta Horti Gotoburg. VII.
1932.
11 Beifräge zur Kenntnis der Embrvologie der Goodeniaceen. — Ibid. XII. 1937.
12. — Notes on the embryology of Globularia vulgaris L. — Bot. Notiser. 1940.
13. Samuelsson, G. Studien liber die Entwicklungsgeschichte der Bliiten einiger
Bicornestypen. — Sv. Bot. Tidskr. 7. 1913.
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