- Project Runeberg -  Den Norske Nordhavs-expedition 1876-1878 / The Norwegian North-Atlantic Expedition 1876-1878 / 4. Bind /
49

(1880-1901) [MARC]
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49

Mundskiven og Tentaklerne have samme histologiske
Bygning som Kroppen. Omkring Mumlen samle Kroppens
transverselle Muskelfibre sig og danne en Slags
Slutte-muskel; fbrovrigt er Muskellagene saavel i Mundskiven som
i Tentaklerne Fortsættelse af Kroppens.

Svælget er meget langstrakt; hos en Polyp, hvis Krop
var 44""" lang. var det 23""" langt; det er cylindrisk, meget
stærkt foldet paatvers, Fig. 45, a, og afdelt i 8 Felter ved
de 8 Septa, som fæste sig paa det, Fig. 45. b. 47, a. Dets
udvendige Flade er beklædt med et Epithel, der bestaar af
flere Lag runde Celler, som have en tynd. klar Membran,
en temmelig stor, excentrisk Kjerne, omgiven af
Protoplasmakorn. Cellerne ere 0,011""" og Kjernen 0,002""",
fuldkommen lig dem, som findes paa Polypkroppens indre
Væg, Fig. 43. b. 46. a. 50, e. Imellem disse
Entoderm-celler sees hist og her lignende runde Celler, men hvori
findes intens brune Pigmentkorn, Fig. 46, b. 50, f.
Indenfor Epithellaget er et temmelig smalt, fibrillært
Binde-vævslag, Fig. 43, c. 51, 6, hvoi’i sees dels Bindevanslegemer,
dels en Mængde fine Saftkanaler, der ere forsynede med
Epithel, hvis lidt aflange Celler ganske udfylde Lumen et.
Fra dette Bindevæv udsendes pyramideformet tilspidsede
Forlængelser, der danne Grundlaget for Svælgets Folder,
Fig. 43, il. 51, c. Paa disse Bindovævspyramider ere
Længde-musklerne fartede, Fig. 43, e. 51, il, imedens Tvermusklerne
vise sig langs deres Grunddel, Fig. 51, e. Hele Svælgets
indre Væg, der er stærkt foldet paatvers og farvet
mørkebrunt, har et Epithelovortræk, der er fæstet til Muskellaget,
og som omtrent i den øverste Halvdel bestaar af polwvdriske
Celler, der have et mørkt, kastaniebrunt Pigmentindhold
og ere lig dem, som ere beskrevne paa Kroppens ydre
Flade. Imellem disse Celler sees hist og her aflange, næsten
klare Celler med en stor. aflang Kjerne og et klart kornet
Indhold, hvilke minde om de encellede Slimkjertler paa
Polyp kroppens og Stokcoenenchymets ydre Flade. .1
Svælgets nederste Halvdel optræder et noget forskjelligt
Epithel-lag. som nærmer sig temmelig meget til det, der beklæder
dets vdre V;eg og Kammervæggene. Det bestaar af runde
Celler. 0,008""" store, som have en tynd, gjennemsigtig
Membran, en rund, noget excentrisk Kjerne, 0,002""" og et
finkornet Indhold, hvori findes mange, temmelig intens
mørkebrune Pigmentkorn, Fig. 48, a. Imellem disse Celler sees
andre, lidt aflange Celler, der ere aldeles klare, have en
rund Kjerne, omgiven af en yderst finkornet Masse og ere
sandsynligvis encellede Slimkjertler, Fig. 48, b, og endelig
iagttages overalt i Svælgets Epithel lange, gjennemsigtige,
brunlige Legemer, i hvis Midte løber paalangs en
spiral-forniig Traad (Neinatocyster), Fig. 51,/. Imellem Folderne
paa Svælget saa es Rester af Molusklarver, Crustacelarver,
Echinodermlarver, samt en hel Del Diatoméer. Alt
Nærings-stofl’e. der vare vel indhyllede i en rig Slimmassé.

Dun norsk«’ Nonlhavsexpeilition. Dmiiolssen og Koren. Pcnnatitlhla.

The oral disk, and the tentacles, have the same
histological structure as the body. The transversal muscular
fibres of the body collect around the oral aperture, and
form a kind of sphincter. The layers of muscles are.
otherwise, both in the oral disk and the tentacles, a
continuation of those of the body.

The gullet is very elongated. In one polyp whose
body measured 44’"’" long, the gullet was 23""" long. It is
cylindrical; very much folded transversally (fig. 45. a), and
is divided into 8 divisions, by the 8 septa which adhere
to it (figs. 45, b. 47, a). It’s exterior surface is covered
with an epithelium, consisting of several layers of round
cells, with a thin clear membrane, and containing, a rather
large eccentric nucleus surrounded by protoplasmic granules.
The cells are 0,011""" wide, and the nucleus is 0,002"""
broad, and exactly like those which are found on the
inner wall of the body of the polyp (figs. 43, b, 46, a, B0, e).
Between these entoderm cells, there are seen, here and
there, similar round cells, but, in which, deep
brown-tinged granules are found (figs. 46, b, 50, /). Inside of the
layer of epithelium, there is a rather narrow layer of
fibrillous connective-tissue (figs. 43. c, 51. b), in which is
seen, partly, connective tissue corpuscles, and, partly, a
multitude of minute secretory ducts; these are furnished
with epithelium, whose slightly oblong cells quite fill out
the channel. From this connective-tissue, pyramidal
acuminated prolongations issue, and these form the
foundation of the gullet folds (figs. 43, d, 51, r). The
longitudinal muscles are adherent to these connective-tissue
pyramids (figs. 43. e, 51, d). whilst, the transversal muscles
appear along their basal part. (fig. 51, e). The inner wall
of the whole of the gullet is much folded, transversely,
and is coloured dark brown. Tt has an epithelium covering,
which is secured to the muscular layer, and which, for
about it’s superior half portion, consists of polyhedrous cells,
having a dark, chestnut brown coloured contents, and. which
are like those which have been described as pertaining to the
body’s exterior surface. Between these cells, there are here
and there seen, oblong, almost clear cells, with a large
oblong nucleus and a. clear granular contents, which recall
to mind the single-celled mucous glands on the body of
the polyp and the exterior surface of the sarcosoma. In
the gullet’s inferior half part, a somewhat different layer
of epithelium appears, and this approaches, pretty much, in
similarity, to that, covering it’s outer wall, and the chamber
walls. It consists of round cells, 0,00S""" wide, with a thin,
transparent membrane, which contain a round, somewhat
eccentric nucleus 0,002""" broad, and a minute granular
contents, in which is seen, many rather deep-brown coloured
granules (fig. 48. a). Between these cells, other, somewhat
oblong cells, are seen; these are perfectly clear, and
contain a round nucleus surrounded by an extremely
minute-grained substance, and are. presumably, single-celled
mucous glands (fig. 48, b); and finally, in the epithelium
of the gullet, there is, everywhere, observed, long, brownish
translucent bodies, in whose middle there runs,
longitudinally, a spiral formed filament (nematocyst) (fig. 51,/).

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