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857

(1914) [MARC] Author: Olof Hammarsten Translator: John Alfred Mandel With: Gustaf Hedin - Tema: Chemistry
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GASES OF THE LYMPH AND .SECRETIONS. 857
animal, is explained by Zuntz ’
by the formation of acid in the tissues,
ami especially in the lymphatic glands, immediately after death, and
this acid in pari decomposes the alkali carbonates < f the Lymph.
The secretions, with the exception of the saliva, in which Pfluger
and Kulz found respectively 0.6 per cent and 1 per cent oxygen, are
almost free from oxygen. The quantity of nitrogen is the same as in blood,
and the chief mass of the gases consists of carbon dioxide. The quantity
of this gas is chiefly dependent upon the reaction, i.e., upon the quan-
tity of alkali. This follows from the analyses of Pfluger. He found
19 per cent carbon dioxide removable by the air-pump and 54 per cent
firmly combined carbon dioxide in a strongly alkaline bile, but, on the
contrary, G.6 per cent carbon dioxide removable by the air-pump and
0.8 per cent firmly combined carbon dioxide in a neutral bile. Alkaline
saliva is also very rich in carbon dioxide. As average for two analyses
made by Pfluger of the submaxillary saliva of a clog we have 27.5 per
cent carbon dioxide removable by the air-pump and 47.4 per cent chem-
ically combined carbon dioxide, making a total of 74.9 per cent. Kulz 2
found a maximum of 65.78 per cent carbon dioxide for the parotid saliva,
of which 3.31 per cent was removable by the air-pump and 62.7 per
cent was firmly combined. From these and other reports as to the
quantity of carbon dioxide removable by the air-pump and chemically
combined in the alkaline secretions it follows that bodies occur in them,
although not in appreciable quantities, which are analogous to the pro-
tein bodies of the blood-serum and which act like weak acids.
The acid or at any rate non-alkaline secretions, urine and milk, con-
tain, on the contrary, considerably less carbon dioxide, which is almost
all removable by the air-pump, and a part seems to be loosely combined
with the sodium phosphate. The figures found by Pfluger for the
total quantity of carbon dioxide in milk and urine are 10 and 18.1-19.7
per cent respectively.
Ewald 3
made investigations on the quantity of gas in pathological
transudates. He found only traces, or at least only very insignificant
quantities of oxygen in these fluids. The quantity of nitrogen was about
the same as in blood; that of carbon dioxide was greater than in the
lymph (of dogs), and in certain cases even greater than in the blood after
asphyxiation (dog’s blood). The tension of the carbon dioxide was
greater than in venous blood. In exudates the quantity of carbon
dioxide, especially that firmly combined, increases with the age of the
1 Buchner, Arbeiten aus der physiol. Anstalt zu Leipzig, 1876; Zuntz, 1. c, 85.
2
Pfluger, Pfltiger’a Arch., 1 and 2; Kitlz, Zcitschr. f. Biologie, 23. It seems as if
Kiilz’s results were not calculated at 760 millimeters Hg. but rather at 1 meter.
3
C. A. Ewald, Arch. f. (Anat. u.) Physiol., 1873 and 1876.

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