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216

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SVENSK BOTANISK TIDSKRIFT. 1919. Bb. 13, H. 2.

NOTES ON THE EMBRYOLOGY OF
SAT-SOTEA SUI
BY
LARS GUNNAR ROMELL

In Chenopodiaceae, the normal type of embryo sac seems to be
a rule without exception, as far as yet known.

However, the embryo sac of Salsola being not hitherto described,
my esteemed preceptor, Prof. O. ROSENBERG, has thought it to be
of some interest to have a species also of that genus investigated.
Mr ROSENBERG also kindly provided me the Zenker- and Flemming-
fixed material. Haidenhains iron-haematoxyline was used for
staining.

Salsola is not a very fit material for embryological work, the sing-
le curved ovule showing in early stages no fixed position relative
to any marked plane. In fact, the trained eye of my friend, Fil.
Lic. G. TàckHorw, but recently discovered that sufficient informa-
tion was to be gathered from my several hundred slides from 1913.

The pollen development could not be followed because the fixing
was not good for that purpose.

In following the development of the embryo sac of Salsola Kali
from the beginning, we find a single sporogenous cell within a nu-
cellar wall, at this stage 2 (or 3) cell.layers thick (see the figure 1 a).
There is formed a tetrade, of which I have seen the chalazal mega-
spore grow out. If that is always so, I am not able to say. While
the embryo. sac is formed in the normal way by the usual divi-
sions (fig. 1 b, c), the nucellus wall grows thicker, so that the ripe
embryo sae lies c:a 6 cell-layers deep.

There are two integuments, as stated by HEGELMAIER and MEU-
NIER, in opposition to VAN TIEGHEM (p. 1406). Both are gene-
rally visible on the dyade stage.

The later stages are described by HEGELMAIER.

The ripe seed of Salsola, as is well known, has an exceedingly
curved embryo, which as a heliciform rolled body fills the whole

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