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medical sciences. 57:5
1. Broman (born 1868), who has published numerous works on embryological
and histological subjects, especially on the structure of the sperms and the
development of the organs of the fetus; Broman teaches macroscopic
Anatomy principally, Fiirst Histology.
527
Allvar Gullstrand.
Physiology, which in Sweden as elsewhere was in earlier times united as a
subject of study at the universities with Anatomy (q. v.), had an independent
professorship allotted to it first in Uppsala, where in 1864 Frithiof Holmgren
(1831—97) became professor of the subject and established the first
Physiological Laboratory in Sweden. Holmgren is best known for his works on
colourblindness and its importance for navigation and railway traffic, and also for his
method of detecting colour-blindness, which is still the one most in use-
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