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Why no description
of this sub-family is
worked oui.
Deficiencies in thc
description of the
genus Philomedes.
Number of genera.
Reproduction of
Pseudo philomedes,
Pleoschisma. and
Tetragonodon.
Reproduction of
Philomedes.
Sub-Family Philomedinae.
S u b - F a m. Philomedinae (part.), G. W. MÜLLER, 1912, p. 24.
Diagnosis: ■— Cf. G. W. Müller, loc. cit.
Remarks: — On account of the comparatively small number of species of this sub-family
that I have had an opportunity of investigating closely, it has not seemed convenient to me
to give in this connection a more detailed description of the sub-family than the one quoted
above, worked out by G. W. MÜLLER. Such a description would, in any case, be very uncertain
because of the uncertainty and incompleteness of the diagnoses and descriptions of the forms
hitherto given.
A natural conséquence of this is that several of the characters that in the present treatise
are included in the description of the genus Philomedes are certain tobe characters of the sub-family.
Five genera of this sub-family have so far been established viz.:
Philomedes, W. LlLLJEBORG, 1853,
Pleoschisma, G. S. BRADY, 1890,
Pseudophilomedes, G. W. Müller, 1894,
Tetragonodon, G. S. BRADY and A. M. NORMAN, 1890.
Paramekodon, ,, ,, ,, ,, ,, ,, ,, ,,
In his synoptic work of 1912 G. W. Müller approves of only two of these five genera,
viz. Philomedes and P seudophilomedes. The genera Pleoschisma and Tetragonodon are in this
work included under the genus Philomedes; Paramekodon is identified with P seudophilomedes.
Unfortunately the descriptions of the species that are included in the genera Pleoschisma
and Tetragonodon are very incomplete and presumably partly incorrect. It seems to me rather
probable, however, that this procedure of G. W. Müller’s is to be considered prématuré at
least in one point. As far as I can see at least the species included under Pleoschisma represent
so different a type that they must be dealt with as a special genus. With regard to Tetragonodon
it does not seem impossible to me that it must be regardée! as a special unit, perhaps as a
subgenus of the genus Philomedes. These questions can, however, only be decitled after a renewed
investigation of these forms. — In the identification of Paramekodon with P seudophilomedes
Müller certainly is correct.
Oecology of reproduction: — With regard to the phenomena connected with the
reproduction nothing at ail is known about the species described under the naines of P seudophilomedes,
Pleoschisma and Tetragonodon.
The reproductive oecology of the genus Philomedes (sensu meo) lias some very interesting
peculiarities to show.
In the following exposition of some of these phenomena in the last-mentioned genus
attention will chiefiy be paid to a single species, Pli. (Pli.) globosa (W. LlLLJEBORG). This
seemed convenient to me partly because of the great part this species has played in the investi-
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