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366
HENNING HORN AF RANTZIEN
The first requirement, that of complete substitution, however, is
möre uncertain in this case. The map (Fig. 3) seems to indicate that
Ihese two forms have different areas of distribution. Vet il should be
pointed out that our knowledge of the detailed distribution of Piil.
arenarium is too fragmentary to provide sure conclusions at present.
However, perhaps we had heiter for the linie being consider this
Aegean form as a subspecies.
vierhapper (I.e.) further pointed out that ssp. aegaeum in a higher
degree than ssp. eu-arenarium is difficult to distinguish from Phleum
crypsoides, Phl. Boissieri and Piil. subulatiim v. ciliatum. However, we
do not think that the form and ciliation of the glumes are very useful
in these cases as being subjects to great variation — Vierhapper lavs
stress lipon these characters — ön the contrary an examination of the
glumella will give decisive results in most cases. Phl. arenarium ssp.
aegaeum has a 3-nerved glumella, Phl. crypsoides a 1-nerved one
(ner-vation very faint and vague) and Phl. siibulatum v. ciliatum has a
5-nerved glumella. Phl. Boissieri is said to have a 3-nerved glumella
(e.g. Boissier 1884, p. 480, Post 1933, p. 718); all material of the
species which the present author has examined shows however a
5-nerved glumella. Perhaps that is here the same multiformity in the
number of nerves in glumella as in Phl. pratense (3—5 nerves) and
Phl. exaratum Griseb. (3—7 nerves) according to my own
examinations. lf |here should exceptionàlly be the same number of nerves (31
in ssp. aegaeum and Phl. Boissieri, the glumella of the former is
± shortly and densely pilose, bul that of the latter glabrous or
sub-glabrous.
For the rest taxonomists have taken two different ways
concer-liing the comprehension of Phl. arenarium.
One way is represented by Fiori and Paoletti (1896—98, p. 58)
and låter bv Fiori (1923, p. 89), these authors bringing in under Phl.
arenarium not onlv the form which Linnaeus described under that
name hut also Phl. exaratum Griseb. (Phl. graecum Hoiss, et Heldr.)
and Phl. crypsoides (Urv.) Hack. ssp. sardoum (Hack.). II is lo be noted
that the same authors onlv lake iniö consideration such forms as are
represented in Italy.
Most o|ber authors, treating from a floristical point of view similar
districts (e.g. Halaczy 1904 and Post 1933), have gone in for an
opposite tendencv, in most cases maintaining all of the earlier
described species. However, no explanation of t lie reasons for the one or the
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