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establishing five sections viz, Euclaytonia, Limnia, Alsinastrum, Naiocrene and
Montiastrum. Characteristic of Euclaytonia is the relatively large, primary, persisting root; in
Limnia the roots are fibrous, and the species mostly annual. Then in Alsinastrum we
have the peculiar Cl. Chamissonis with all the leaves opposite, and with bulbiferous
stolons; in Naiocrene which is, also, perennial, bulblets are developed in the axils of many
of the cauline leaves. Finally in Montiastrum the species are strictly annual with mostly
alternate leaves, and with mostly two of the stamens suppressed. In other words several
of the species are annual, and among those which are perennial we meet with some that
possess large persisting roots, and others in which these are fibrous and fugacious, but
where the vegetative reproduction is secured by means of bulblets, aerial or subterranean.
Now in respect to the structure of the shoot, we have shown[1] that the
inflorescence is frequently lateral, and indeed most frequently so. The inflorescence is of the
cymose type, but the lateral ramifications are frequently monochasia of the type cicinnus
or scorpioid cyme, as described by Wydler[2]. Preceding the inflorescence we frequently
notice one pair of opposite leaves, which by Eichler and Wydler have been defined as
prophylla; the flowers themselves are mostly destitute of such foreleaves, but there are
some species, however, in which one of these is developed, especially in the lower part
of the inflorescence. Most frequently the two foreleaves, preceding the inflorescence,
are the only leaves of the aerial stems, but in some few species, as will be shown later,
the stems are leafy from the base to the inflorescence, opposite in Cl. Chamissonis,
alternate in Cl. linearis and parvifolia.
In outlining the genus Gray (l. c.) did not distinguish between species with the shoot
terminated by an inflorescence or being purely vegetative, nor did he consider the
character “annual” or “perennial” of much importance to his classification. Nevertheless
in all the members of Montiastrum the shoot is terminated by an inflorescence, but not
so in any of those of Limnia. The duration of life of these species, annual or perennial,
does not seem to be of any importance, inasmuch as we know that annuals sometimes
become perennial, and in Claytonia this is illustrated by Cl. Sibirica according to Gray
(l. c).
Let us now examine the types of the genus, classified as suggested by Gray, and
considered inseparable from the genus Claytonia.
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