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cotyledons above ground; at this stage two leaves, succeeding the cotyledons, are already
visible, and the plant is now ready to meet the first winter. During the winter the
hypocotyl shows a gradual wrinkling, by which the plumule becomes pulled down beneath
the surface of the ground, and the root continues its growth vertically, and to a very
considerable depth. In the following spring several leaves develop and form soon a
small rosette, while increase in thickness commences in the hypocotyl and in the basal
portion of the root. A fully matured specimen is figured on Plate 1, showing the large
root, the rosette of leaves, and the inflorescences, all axillary. In this illustration we
have before us the picture of a plant that inhabits the highest peaks of the Rocky
Mountains. And at the stage figured the leaves and inflorescences for the coming year are
already developed, and visible when we lay a longitudinal section through the rosette
and the apical portion of the hypocotyl. The leaves vary from spatulate to
dilatedcuneate or obovate (figs. 3—5); the inflorescence (fig. 2) is cymose with the two large
foreleaves empty, and with only one bract to each cyme. In respect to the structure
of the flower (figs. 6 and 8) this agrees with Claytonia proper; so does also the opening
of the capsule (figs. 9—14), and the seed (fig. 15).
If we compare this species of Claytonia with the other high alpine plants of the Rocky
Mountains, the deep-seated root seems somewhat anomalous; but otherwise the
leaf-structure, for instance, is that of many alpine species, this being isolateral, the stomata
being distributed over both faces of the blade, and the chlorenchyma representing a
palisade-tissue throughout.
A similar development of the root and foliage exists in C. arctica Adams and C.
tuberosa Pall., but not in the other species pertaining to this section.
Claytonia Virginica L. (Plate II fig. 1).
This is, also, a relatively large-rooted species, in which the primary root increases
considerably in thickness, but only the basal portion, since the slender apical part dies
off entirely. In respect to the seedling-stage we find only one cotyledon developed,
hence reminding of Ficaria, Cyclamen, Dicenira etc. The ramification of the shoot is
monopodia!, but the leaves do not persist throughout the winter as is the case with C.
megarrhiza Parry. In regard to the leaf-structure this is dorsiventral so far as concerns
the chlorenchyma, but not in respect to the stomata, which occur on both faces of the
blade. Claytonia Caroliniana Michx. and C. lanceolata Pursh agree with C.
Virginica L.
Claytonia sarmentosa C. A. Mey.
The last member of Euclaytonia, since we prefer to place C. asarifolia Bong, under
the section Limnia. In C. sarmentosa C. A. Mey. (Plate II fig. 2) we notice the deep
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